| Smelowskia johnsonii G.A. Mulligan | |
| Publ. & Syn. | Smelowskia spathulatifolia Velichkin, Bot. Zhurn. 59: 1805 (1974). - Nomen novum for Hutchinsia calycina var. integrifolia Seem., Bot. Voy. Herald: 25 (1852). Described from the vicinity of Cape Krusenstern [perhaps Cape Mulgrave], Kotzebue Sound (Alaska). - Smelowskia calycina var. integrifolia (Seem.) Rollins, Rhodora 40: 300 (1938). - Smelowskia calycina subsp. integrifolia (Seem.) Hultén, Acta Univ. Lund., n. s., sect. 2, 41, 1: 869 (1945) [comb. illeg., without basionym reference]. |
| Notes | Yurtsev: Melanidion boreale sharply contrasts to species of true Smelowskia, e.g., in the monopodial structure of rosette and the ternate leaves reminding those of Ermania parryoides. Mulligan, Elven, and Murray: Appel and Al-Shehbaz (2003) wrote about Smelowskia: "One of its segregates, Melanidion, perhaps merits recognition". Mulligan commented that he does not agree with this. In his opinion, the differences are not large enough for recognition of Melanidion as a genus. Melanidion nested within Smelowskia in the analysis of Warwick et al. (2003a) and were synonymized with it by Al-Shehbaz and Warwick (2006b). However, in microsatellite data, the three North American Melanidion species (S. borealis, S. ovalis, S. pyriformis), together with the non-arctic northeastern Asian S. inopinata, constituted a sister to Smelowskia s. str. (Carlsen et al. 2010). The matter may have to be re-evaluated when more material of the other previous segregate genera (Ermania, Gorodkovia, Hedinia) is included in the analysis. |
| Chromosomes | (1) 12 (2x, x = 6). - Alaska. - Murray and Kelso (1997, for S. calycina var. integrifolia). (2) 18 (2x?, x = 9?). - Far East (N). - Zhukova and Petrovsky (1984, for S. porsildii); Zhukova (unpubl.). (3) 22 (2x?, x = 11?). - Alaska. - Johnson and Packer (1968, for S. calycina var. integrifolia); Dawe and Murray (1979, for var. porsildii); Rollins (1993). (4) 24 (4x, x = 6). - Far East (N). - Yurtsev and Zhukova (1972, for S. porsildii); Yurtsev et al. (1975). (5) 32 (4x?, x = 8?). - Far East (N). - Zhukova and Petrovsky (1984, for S. porsildii). |
| Geography | American Beringian: ALA. |
| Distribution | D = ? E = R AW = r CW = r [ key ] |
| Parent taxon | Smelowskia C.A. Mey. |
| PAF ID | 670205 |