Salix sphenophylla A.K. Skvortsov
Publ. & Syn.A.K. Skvortsov, Fl. Arct. URSS 5: 62 (1966). - Nomen novum for Salix cuneata Turcz. ex Ledeb., Fl. Ross. 3, 2: 623 (1850), non Nutt., N. Amer. Sylv. 1: 66 (1842). Type (LE): Siberia: "inter Jakutsk et Ochotsk", 1835, leg. Turczaninow.
NotesArgus, Elven, Murray, and Skvortsov: The ranges of the two proposed races of Salix sphenophylla are not clear. Skvortsov (1966) reported subsp. sphenophylla from Siberia (Anabar-Olenyok, Kharaulakh, Yana-Kolyma) and as questionable from Chukotka and western Alaska. He reported subsp. pseudotorulosa from Chukotka and western and northern Alaska. Hultén (1968a) mapped subsp. pseudotorulosa (as S. arctica subsp. torulosa) as Alaskan and northern Cordilleran. There is an overlap and reports of intermediates in Chukotka but Yurtsev commented that the major part of the collections from Chukotka well fits subsp. sphenophylla. Murray and Elven have compared specimens (ALA) from Chukotka and Alaska and find no consistent difference. Our current proposal is of a northern Siberian subsp. sphenophylla and an amphi-Beringian subsp. pseudotorulosa. We have sorted chromosome number reports accordingly. Skvortsov commented that subsp. pseudotorulosa could be an ecotype of S. arctica but the parapatric ranges count against that interpretation.
       There is another possibility, suggested by Hultén (1968b): "It seems possible that this taxon i.e., subsp. [pseudotorulosa] represents the introgression between S. sphenophylla A. Skvorts. (S. cuneata Turcz. non Nutt.) with S. arctica". . Salix sphenophylla is reported to be diploid, whereas S. arctica is polyploid. Hybridization is probable where they meet and intermediate ploidy levels and morphology might be expected. Subspecies pseudotorulosa should be reexamined with this in view. However, nearly all chromosome reports of S. sphenophylla from Chukotka are of diploids.
Chromosomes(1) 76 (4x). - Far East (N), Alaska, Canada, U.S.A., Greenland. - Several reports, some Russian ones for subsp. crassijulis and subsp. jamu-taridensis.
(2) >100 114 (6x). - Far East (N), Alaska, Greenland? - Several reports.
(3) ca. 120. - Asia (Altai). - Sokolovskaya and Strelkova (1948a).
(4) 190 (10x). - Europe (Iceland). - Löve and Löve (1956b, for S. callicarpaea, see comment under S. glauca above).
Löve and Löve (1975a) assigned the reports of tetraploids by Holmen (1952), Mosquin and Hayley (1966), Johnson and Packer (1968), Suda and Argus (1969a), and Löve et al. (1971) to S. arctica s. str., those of hexaploids by Suda and Argus (1969a) and Zhukova et al. (1973) to S. crassijulis, and they neglected the other reports and numbers. However, also the Löves' reports of hexaploids for S. "cordifolia" from Iceland and Greenland (Löve and Löve 1975c) and of decaploids for S. callicarpaea from Iceland (Löve and Löve 1956b) should refer to what now is considered S. arctica. There seems to be evidence for at least two main ploidy levels in S. arctica s. lat., widespread tetraploids (and perhaps higher numbers) and perhaps Beringian hexaploids.
GeographyAsian (N/C) - amphi-Beringian - Cordilleran.
Parent taxonSalix L.
Child taxa Salix sphenophylla subsp. pseudotorulosa A.K. Skvortsov
Salix sphenophylla subsp. sphenophylla
PAF ID580218
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Panarctic Flora Editor-in-Chief: Reidar Elven (Natural History Museum, University of Oslo)
Editorial Committee: Reidar Elven, David F. Murray (Museum of the North, University of Alaska), Volodya Yu. Razzhivin (Komarov Botanical Institute, Russian Academy of Sciences), Boris A. Yurtsev [deceased] (Komarov Botanical Institute, Russian Academy of Sciences)